Swifty (Jeremy Renner) is a little fox with big dreams. Working in the the mailroom of the Arctic Blast Delivery Service, he dreams of one day becoming a Top Dog (the Arctic's star husky couriers and town heroes). Tired of waiting, he decides to prove himself worthy. Swifty steals a sled to make his first delivery - a mysterious package to a secret location - where he stumbles upon the hidden fortress of evil genius Otto Von Walrus (John Cleese) and his loyal army of oddly polite puffin henchmen. When Swifty discovers Otto’s secret plan to drill beneath the Arctic surface and unleash enough ancient gas to melt the Arctic, he enlists the help of his friends: PB, an introverted polar bear (Alec Baldwin), Lemmy, a scatterbrained albatross (James Franco), Magda, his boss (Anjelica Huston) and Jade Fox (Heidi Klum), a tough as nails mechanic. This ragtag group of Arctic misfits must to band together and stop Otto Von Walrus' sinister plan to become the world's supreme leader and save the planet – before it’s too late!